Science in a Troubling Political Climate – Dr. Chris Widga – Part 2

New Hampshire doesn’t have a state museum.  I never realized there were such things until someone I interviewed mentioned a mastodon fossil in Albany, which prompted me to travel to the NY State Museum soon after to see it.

Not understanding what a state museum is, it shocked me that there was no admission fee; anyone from anywhere could visit the museum at no cost.  I marveled then—as I marvel still—that such places exist. (To be clear: not all state museums are free.)

The Cohoes Mastodon at the NY State Museum in Albany, NY; picture taken by the author of this blog.  I learned of this mastodon thanks to Dartmouth professor, Dr. Roger Sloboda, after interviewing him for a piece I was writing about a mammoth & mastodon exhibit at the Boston Museum of Science in 2012.

 

 

Illinois not only has a state museum, that museum is made up of five separate museums with over 13 million artifacts.  And in 2015, Governor Bruce Rauner wanted to close it completely.

During a messy and contentious budget battle, the museum was shuttered for nine months, only to be reopened this past July with a new $5 admission fee.  But by then, most of the staff had gone, forced to take jobs in other places as their future at the museum was decidedly uncertain.

 

Screenshot from this page of the Illinois State Museum website.

 

No one knows this better than Dr. Chris Widga, who had been a vertebrate paleontologist employed at the Illinois State Museum.  He now works at the Center for Excellence in Paleontology at East Tennessee State University (ETSU).

“The whole question of Channel Islands and island mammoths probably got me through last winter,” Dr. Widga explained as we spoke by phone.

We were discussing the effect islands had on proboscidean evolution and the exciting recent research done in part by researchers from The Mammoth Site and the National Park Service.

“In Illinois, as the State government was falling apart around my ears, as the State Museum was closed, I basically closed my door and was doing the analysis for the Quaternary International article. In so doing, I was thinking about these pygmy mammoths. As it’s icy outside and subzero for about six weeks at a time, that kept my sanity.” He laughed.  “So the Channel Islands has been my refuge, I guess, even though I’ve never actually been out on one of them.”

The move from Illinois to Tennessee was not just a contrast in physical environments.  It also meant moving from a scientific institution founded in 1855 to one that has been open for just 10 years.  Dr. Widga explained that a mere two weeks prior to his start date at ETSU, the university formed a partnership with a local science center.   The ETSU staff maintains the collections, conducts research, and oversees excavations at the nearby Gray Fossil Site.  The science center staff is responsible for educational activities within the museum and overall maintenance.

“Their [educational] philosophy is very similar to ours,” Dr. Widga said of the science center. “It’s inquiry-based. We want people to come in and learn through asking questions rather than just be spoon-fed facts.”

So much of what Dr. Widga has done involves public outreach.  From videos about the collections at the Illinois State Museum to long-distance learning programs like The Mammoth Expedition, work he did in conjunction with Dr. Katy Smith at Georgia Southern University and with the Milwaukee Public Museum.

When I commented on how much I loved that kind of publicly accessible information, his response was, “Part of that is because I’m in a museum. I’m not buried under coursework and teaching. Outreach is valued. The way you justify your existence in a museum is to connect with the public.  And part of that is figuring out how we can connect with the public in ways where it’s an exponential relationship.”

In other words, not having a one-on-one conversation with a museum visitor, but creating a website about the Ice Age in the Midwest, for example.

Figure from a presentation done by Dr. Chris Widga as part of the National Science Foundation grant received; image courtesy of Chris Widga.

 

Despite everything he’s gone through, there is no question Dr. Widga loves what he does.  It permeates his voice when he speaks of paleontology, and it prompted me to ask if he ever becomes excited at work.  His response was a definitive ‘YES.’

By way of explanation, he quoted his now-retired colleague, Dr. Jeff Saunders, who used to say, “‘Going to work in the morning was like going to Disney Land everyday.’”

Not only did the two scientists literally work across the hall from each other at the Illinois State Museum, they were apparently known to shout out excitedly to the other whenever one read a great article or wanted to share a relevant scientific image.

“Part of the reason I like museums is because you just never know!” Dr. Widga continued. “Some of the new things come from the collections; some of the new things come from new papers. You read them and you’re like, ‘oh, this explains it!’ It was something that you had been working on for a long time and, all of a sudden, somebody else had that last piece of the puzzle that puts the whole thing together.

“At least once a day—even on the worst days—there’s something that comes through and I’m like, ‘oh, this is so cool!’”

Proboscideans at Morrill Hall at the University of Nebraska State Museum of Natural History; image courtesy of Chris Widga

 

The seemingly idyllic work environment in Illinois lasted for a decade until 2015. Despite protests, a MoveOn.org petition and public outcry about the museum closing, Dr. Widga and his colleagues were forced to consider other options.  The fate of the museum was out of their hands.

“There was a point as I started looking for jobs last year that I asked myself, you know, do I want to continue in this vein?”

“I’d watched many of [my colleagues] that had taken jobs in Research One institutions [become] totally burned out.  Or they’d kind of gone in weird and funky directions, not because the research was taking them in that direction, but because they were getting pressure from their institution to go in a certain direction or something like that.

“And that was part of the fun of the Illinois State Museum is that I could work on anything. Nobody was saying, ‘You have to work on elephants.’ That was a choice that was mine. Nobody was saying, ‘well, you have to work on dogs.’ That was a choice that was mine. You could chase whatever questions were out there.

“The feedback that I got from the people that interviewed me [was that] they were very interested in what I did.  It was a very different situation than what we were going through at the Illinois State Museum where, essentially, you were being told, ‘what you do is not important.  And none of what you do—your position, your entire existence—is important.’ [The feedback I got while interviewing for other jobs] revived this idea that what we do is important, and it’s exciting.”

I couldn’t help but compare his experience in Illinois to the general anti-science climate in our government today.  It was particularly interesting for me to speak with Dr. Widga about his paper on Pleistocene ecology a day or so after the House Science Committee’s so-called hearing on climate change.  Dr. Widga’s infectious enthusiasm took a very somber turn, as he conceded how difficult things become when “politics starts really driving the boat and reason takes a back seat.”

“That won’t change any of the science,” he added, “[but] it may change how the science is funded. It also won’t change any outreach that we do or the educational activities! In fact, if anything, it’s going to make those seem more important and put more emphasis on those.

“We can talk about the scientific community writ-large, but certainly within the paleontological community, you will find very few working paleontologists, working scientists, who say that education and outreach is not a good thing anymore.

“It used to be that you could just hole-up and do your research and never really interact with the public.  But if anything, this whole process [with the IL governor and the Illinois State Museum] has made us realize that that can’t happen.

“There’s this realization that pre-dates this modern political atmosphere: That you really do need to work with the public and you need to make sure that the point of what you’re doing is out there. Not just in terms of dinosaurs are always cool so therefore that’s why we’re doing it. But we’re also doing it to learn more about how our world works–the nuts-and-bolts of how ecosystems are put together, the nuts-and-bolts of how climate changes impact those ecosystems–that has real-life implications for today and into the future.

“And there’ve been some really loud voices in the last couple of years that have said that over and over and over again. Some of which are people like Jacquelyn Gill! And that is a big shift in science. It’s a big shift in science communications.

“I’m glad that we were moving on that before the current [political] atmosphere because it makes it much more difficult to sideline us as, you know, a bunch of eggheads.”

It didn’t take long for our conversation to take a positive swing upward, as Dr. Widga then described possible future projects involving scientists across the country.

His statement “I’ve always been of the opinion that science is a collaborative effort” couldn’t be more apt.  And I, for one, cannot wait to see what he and his colleagues work on next.

Artwork by Velizar Simeonovski based on scientific research at Mastodon Lake in Aurora, IL; courtesy of Chris Widga

*****

THANK YOU, Dr. Chris Widga, for your generosity of time and spirit in speaking with me about paleontology and the difficulty you’ve gone through.  I loved conversing with you, and I’m eager to read about, watch or see the projects you dive into next!

 

References:

  1. Illinois State Museum reopens to public after nine-month shutdown, John Reynolds, The State Journal-Register, July 2, 2016
  2. Closing decimates Illinois State Museum management, Chris Dettro, The State Journal-Register, December 27, 2015
  3. Much of Illinois State Museum management leaves amid closure, Chicago-Tribune, December 28, 2015
  4. Museums caught in middle of state budget showdown, Steve Johnson, Chicago-Tribune, June 25, 2015
  5. Rainer prepares to close state museums, shutter some prisons to balance ‘phony’ Democratic budget, Becky Schlikerman, Chicago Sun-Times, June 24, 2015
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Stegodon: Does this ancient elephant have origins in Asia?

So much has been said in recent years about the wealth of fossils in China. Almost all of it about dinosaurs: exciting new species, feathered fossils, nest upon nest of dinosaur eggs.  There is no doubt that China holds exciting clues to the history of our planet; one has only to wait to hear of the next discovery.

Within the past few months, yet another exciting find was revealed, but this time about a little known mammalian ancestor: Stegodon.

 

Stegodon by artist Hannah Stephens

Painting of a Stegodon by artist Hannah Stephenshannahleestudio.com.

 

The name Stegodon, to me, evokes ‘dinosaur’, not ‘mammal,’ but this was, indeed, an ancient animal.  Its fossils resemble those of other similar mammals, from mastodons to mammoths to today’s elephants.

 

 

Alexandra van der Geer - shrinking elephants

Figure 1: Reconstruction of four insular dwarf proboscideans with their respective mainland ancestors. Mainland proboscideans: 1, Palaeoloxodon antiquus; 2, Mammuthus columbi; 3, Stegodon zdanskyi [stegodon found in China]. Insular proboscideans: 4, Palaeoloxodon ‘mnaidriensis’; 5, Palaeoloxodon falconeri; 6, Mammuthus exilis; 7, Stegodon aurorae [a type of dwarf stegodon found in Japan]. Based on skeletons at Museo di Paleontología, University of Rome, Italy (1), American Museum of Natural History, New York (2), Taylor Made Fossils, U.S. (3), Museo di Paleontología e Geología G.G. Gemmellaro, Palermo, Italy (4), Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt, Germany (5), Santa Barbara Museum of Natural History, Santa Barbara, U.S. (6), Taga Town Museum, Honshu, Japan (7). Photos 1–2, 4–7 George Lyras, photo 3 courtesy of TaylorMadeFossils.com, reproduced here with permission.

From The effect of area and isolation on insular dwarf proboscidea by Alexandra A. E. van der Geer et al; photo and caption courtesy of Dr. Alexandra van der Geer.

 

When Stegodon skulls with tusks attached have been found, many (but not all) of the tusks are close together–preventing the trunk (the ‘proboscis,’ from which this group gets its name; proboscis —> proboscidea) from hanging between them.

They lived in what is now Africa and Asia, causing continued debate over its place of origin. Until recently, the oldest known Stegodon fossil, a 6.5+ million-year-old partial molar from Kenya, was described by William J. Sanders in 1999.  That record changed this past December when Dr. Hong Ao and his colleagues published their results dating the sediment in the Lanzhou Basin, China, from which a number of fossils–including that of a Stegodon–were found.

And that Stegodon was found to be between 8 – 11 million years old.

 

GSA Geologic Time Scale - Neogene

Detail of the Geologic Time Scale, created by the Geological Society of America.  Stegodon is believed to have existed between the Miocene to the Pleistocene, a relatively small segment of time in Earth’s overall history, but still considerably longer than that of our own species!  (You can view the time scale in much better detail here.) 

 

The fossils of the Stegodon, along with at least 5 other species, were actually found in the 1980s by Professor Xing Zhang of Northwest University in China.  Given the length of time between the fossil excavations and the recent dating of these fossils, one might wonder why determining the fossil age took so long.

Dr. Ao, a scientist at the State Key Laboratory of Loess and Quaternary Geology, Institute of Earth Environment, Chinese Academy of Sciences, and his colleagues state that western China does not have suitable material for radiometric dating, an oft-used method for this purpose.

When I asked why this was so, Dr. Ao replied by email, “Because volcanic activities are rare in western China during the late Cenozoic, it is difficult to find  in situ tephros or tuffs for radiometric dating (e.g., 40Ar/39Ar dating).”

Instead, they conducted a magnetostratigraphic study, one in which they determined the age of the rock through the polar reversal record.  Combining this with analysis of the fossils provided evidence that this Stegodon is 2 – 5 million years older than that of the Kenyan partial molar.

“We are indeed surprised by our dating results,” Dr. Ao continued, “which document that the Lanzhou Stegodon is the oldest Stegodon worldwide, although the Stegodon  fossil [was] not discovered by us. However, our dating results document it to be the oldest known Stegodon fossil.”

Dr. Hong Ao 1

 

Dr. Hong Ao 2

Dr. Hong Ao 3

Dr. Hong Ao 4

Images of Professor Yongxiang Li (from Northwest University) and his master student as well as several employed workers who helped to excavate mammal fossils, Lanzhou Basin, China; photos courtesy of Dr. Hong Ao.

 

indricotherine fossil

indricotherine fossil2

indricotherine fossil3

Images of indricotherine fossils found in Lanzhou Basin, China; photos courtesy of Dr. Hong Ao.

 

Dr. Ao himself has been working with fossils in the Lanzhou Basin for 5 years.  When asked how  he and his co-authors chose to work together on this recent paper, he wrote, “I have collaborated with them on broad subjects before, thus I [invited] them to join [in] this research.”

Finding information about this extinct species is difficult.  Unlike mammoths or mastodons, Stegodon does not appear to be a popular ancient animal.

Fortunately, Dr. Alexandra van der Geer–paleontologist, indologist, ethno-zoologist and author –has not only studied this species, she very generously made herself available for questions.  Currently, Dr. van der Geer is an Associate Researcher at both the National and Kapodistrian University of Athens and the Naturalis Biodiversity Center in the Netherlands. She is part of the Isolario project, a project that studies biodiversity and cultural evolution within an island context.

When I asked why there was such a dearth of information on Stegodon, she wrote, “We can think of several reasons for this.”

“First of all, stegodons typically are the elephants of Southeast and East Asia, where most countries did not have the resources and opportunities most Western countries had when it comes to scientific research and excavations. These activities are costly, and since they don’t have a direct use in the sense that they don’t advance the medical, technical or economic levels of a country, they have, understandably, a lower priority. Furthermore, because of this region, half the publications (especially before the 1990s or so) you can find are in Chinese or Japanese, which is not very helpful to the [English-speaking] world.

“Secondly, stegodons are forest elephants. Forest areas are very unlikely places for the long-term preservation of organic materials: everything is eaten, digested or otherwise broken down into smaller components in no time. The tropical (warm, humid) climate of these forests is not helpful either, as decomposition is much faster here. Stegodon remains only have a chance to be preserved when (1) they are covered fast, such as with river sediments, volcanic ashes etc., (2) or are in an oxygen-free environment, such as sunken deeply into a swamp, (3) or were deposited in a natural fridge such as limestone caves where they are gradually covered in clayish sediment or [travertines]. The same is valid for Palaeoloxodon, the Old World fossil elephant, but Europe has many limestone caves, which are excellent for preservation (for a nice [travertine]-preserved negative skull, see Stuttgart museum: skull cast SMNS nr. 32888 from Bad Cannstatt).

“As you can deduct from these preservation issues, it is more likely to find molars and tusks than skeletal material, which is much softer. The vast majority of proboscidean findings all over the planet consists of molars and tusks, and that is not for nothing. Inherently this means that there is much more information about their dentition and diet than about their bodies.”

I was interested in understanding why Stegodons are portrayed as hairless animals, so very similar to contemporary elephants.  Was this just an artistic guess?

“The hairlessness of stegodons is not an artistic guess but a scientific guess instead,” Dr. van der Geer answered.  “Very large animals with thick skins (pachyderms) in a (sub)tropical environment are unlikely to have a significant hair covering. Elephants lost their hairs secondarily. The information for hair growth is not lost, and baby elephants still have a thin, woolly coat. Woolly mammoths lived in the cold, temperate zones, and needed hair, so they were covered in a thick layer of hairs, and for this is evidence (mummies preserved in the permafrost), but the other mammoths (M. meridionalis, M. columbi, M. exilis, etc.), [did] not, and it’s generally assumed that they had a light coat fitting to the temperate zones.

“Tropical and subtropical stegodons almost certainly did not have any coat that’s worth mentioning. Stegodons of temperate zones, however, may have been more hairy. Indeed, the lack of hairs makes them look more like today’s elephants.”

 

Alexandra van der Geer - Stegodon ganesa-model-I.Vjdchauhan-SiwalikHills

Photo of the two life-size models of Stegodon ganesa;photo courtesy of Dr. Gerrit van den Bergh (University of Woolongong, Australia); special thanks to Dr. Alexandra van der Geer.

 

“Note, however, that the proboscis is carried very differently. Their tusks are set very close to each other, so the proboscis doesn’t fit in between as in modern elephants, Asian and African alike. This means that the mobility of their proboscis was more restricted, relative to their living relatives.”

 

Alexandra van der Geer - Flores-excavation-31-stegodon-florensis

Fossils of Stegodon florensis insularis, from Flores, Indonesia; photo courtesy of Dr. John de Vos (Naturalis, the Netherlands); special thanks to Dr. Alexandra van der Geer.

 

Alexandra van der Geer - stegodon-timorensis-mandible

Mandible (and holotype!) of  Stegodon timorensis; photo courtesy of Eelco Kruidenier (Naturalis, the Netherlands); special thanks to Dr. Alexandra van der Geer.  Anyone familiar with proboscidean teeth and jaws will recognize the similarities instantly.

 

But how do we know that Stegodon–a rather enormous animal–evolved into something smaller?

“[D]warfs and giants are relative. Something can be a dwarf, yet have a considerable size. When we speak of dwarf stegodons, we mean stegodons that are much smaller than their ancestors. For this, you have of course to have identical or otherwise similar elements from both the descendant and the ancestor in order to compare reliably,” she continued.

“The expectation is that dwarf stegodons must have existed on the islands, following the so-called island rule, according to which large animals get smaller in isolation. There is sound evidence that this rule still stands, and is even more pronounced for fossil species (see Lomolino et al., 2013, in Journal of Biogeography).

“Indeed, the many fossil molars from the Southeast Asian islands (‘Wallacea’) are all much and much smaller than the same molars from their mainland ancestors (see Van den Bergh, 1999). True, you first have to know what is the ancestor, and for this you need information about morphology, or how the molars, tusks, skulls and postcranial elements look like. After that, you compare the sizes.

“Note that if a molar is, for example, half the length of the same molar of its ancestral species, the body weight of that animal must have been a quarter of that of its ancestor! (the cubic law: linear reduction 50% means volume reduction 50% of 50%).”

Alexandra van der Geer - Flores-stegodon-florensis

Molar of Stegodon florensis; photo courtesy of Dr. Gerrit van den Bergh (University of Woolongong, Australia); special thanks to Dr. Alexandra van der Geer.  

 

“The most interesting dwarf stegodon is Stegodon sondaari, named after the Dutch palaeontologist Paul Yves Sondaar (1934-2003), expert in fossil insular mammals. This stegodon lived on the island of Flores about a million years ago, and weighted only about 15% of the weight of its ancestral species, S. elephantoides (see Van der Geer et al., 2016, in Journal of Biogeography, doi:10.1111/jbi.12743).

“Sondaar’s dwarf stegodon is not the smallest stegodon, that honour goes to the Sumba stegodon (S. sumbaensis), of only 8% of the original weight. Sondaar’s stegodon is interesting because it may have witnessed the arrival of early humans, possibly the ancestors of the Hobbit, or Homo floresiensis. Its fossils are contemporaneous with primitive lithic artefacts, dated to about a million years ago (see Brumm et al., 2010, in Nature 464, pp. 748–752).”

 

Alexandra van der Geer - Sumba-stegodon-sompoensis-holotype-in-Naturalis-Leiden-2

Molar (and holotype!) of Stegodon sompoenisphoto courtesy of Dr. Gerrit van den Bergh (University of Woolongong, Australia); special thanks to Dr. Alexandra van der Geer.  

 

“[R]ecently,” she concluded, “one of the island dwarf stegodons (S. timorensis of Timor) has been dated to about 130 thousand years ago (see Louys et al., 2016, in PeerJ 4:e1788). This excludes, according to the authors, an anthropogenic cause for its extinction, because humans had not yet arrived at the island.”

 

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So many people helped with this blog post!  (But please remember that any errors are my own.)

Many, many thanks to Dr. Hong Ao (Dr. Ao Hong) from the State Key Laboratory of Loess and Quaternary Geology (Chinese Academy of Sciences) for his fascinating responses and the great images of fossil excavations in the Lanzhou Basin.  I am thrilled that he was willing to answer questions about his research and that of his colleagues! It was a great honor and a pleasure connecting with him!

I am indebted to Dr. Alexandra Van der Geer, who very kindly (and so very quickly–despite everything else she has going on!!) answered specific questions about Stegodon that I could not find anywhere else and who provided pictures of dwarf Stegodon fossils.  It was an equally great honor and pleasure connecting with her!

A mastodon-sized thank you to the amazing Dr. Katy Smith for providing needed and hard-to-find material on Stegodon fossils!

And an enormous thank you to artist Hannah Stephens for her depiction of a Stegodon as it may have appeared in life.  I am particularly moved by the warmth of its intelligent-looking eyes, and I love the tones within its skin.  I adore this picture.  I am grateful to have it in this post;  I am thrilled to have the actual painting hanging on my wall!  Please be sure to check out her artwork at: http://hannahleestudio.com or http://hstephens.blogspot.com

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References from Dr. Alexandra Van der Geer:

  1. Brumm A, Jensen GM, van den Bergh GD, Morwood MJ, Kurniawan I, Aziz F, Storey M (2010) Hominins on Flores, Indonesia, by one million years ago. Nature 464, 748–752.
  2. Lomolino MV, van der Geer AAE, Lyras GA, Palombo MR, Sax DF, Rozzi R (2013) Of mice and mammoths: generality and antiquity of the island rule. Journal of Biogeography 40, 1427–1439.
  3. Louys J, Price GJ, O’Connor S. (2016) Direct dating of Pleistocene stegodon from Timor Island, East Nusa Tenggara. PeerJ 4:e1788
  4. van den Bergh GD (1999) The Late Neogene elephantoidbearing faunas of Indonesia and their palaeozoogeographic implications; a study of the terrestrial faunal succession of Sulawesi, Flores and Java, including evidence for early hominid dispersal east of Wallace’s line. Scripta Geologica 117, 1–419.
  5. van der Geer AAE, van den Bergh GD, Lyras GA, Prasetyo UW, Due RA, Setiyabudi E, Drinia H (2016) The effect of area and isolation on insular dwarf proboscideans. Journal of Biogeography, doi: 10.111/jbi.12743

References used in this blog post:

  1. New magnetochronology of Late Miocene mammal fauna, NE Tibetan Plateau, China: Mammal migration and paleoenvironments; by Hong Ao, Peng Zhang, Mark J. Dekkers, Andrew P. Roberts, Zhisheng An, Yongxiang Li, Fengyan Lu, Shan Lin, Xingwen Li; Earth and Planetary Science Letters; 1o December 2015
  2. Oldest record of Stegodon (Mammalia: Proboscidea); by William J. Sanders; Journal of Vertebrate Paleontology; Vol. 19, No. 4, Dec. 13, 1999, pp. 793 – 797
  3. Fossil elephantoids, Awash paleolake basins, and the Afar triple junction, Ethiopia; by Jon E. Kalb; Palaeogeography, Palaeoclimatology, Palaeoecology; 1995, pp. 357 – 368
  4. The effect of area and isolation on insular dwarf proboscidea; by Alexandra A. E. van der Geer, Gerrit D. van den Bergh, George A. Lyras, Unggul W. Prasetyo, Rokus Awe Due, Erick Setiyabudi, and Hara Drinia; Journal of Biogeography; 11 March 2016.
  5. Magnetostratigraphy – concepts, definitions, and applications, by Cor G. Langereis, Wout Krijgsman, Giovanni Muttoni, and Manfred Menning; Newsletter on Stratigraphy, Vol. 43/3: 207–233, April 2010
  6. Mammoths and Mastodons of the Ice Age, by Adrian Lister, Firefly Books, 2014
  7. Mammoths, by Adrian Lister and Paul Bahn, University of California Press, 2007
  8. The Proboscidea: Evolution and Palaeoecology of Elephants and Their Relatives, Edited by Jeheskiel Shoshani and Pascal Tassy, Oxford Science Publications, 1996
  • Stegodontidae: evolutionary relationships by Haruo Saegusa, pp. 178 – 190, The Proboscidea: Evolution and Palaeoecology of Elephants and Their Relatives
  • Palaeobiogeography of late Neogene African and Eurasian Elephantoidea by Jon E. Kalb, David J. Froehlich, and Gordon L. Bell, pp. 117 – 123, The Proboscidea: Evolution and Palaeoecology of Elephants and Their Relatives